![[Member-Image]](../assets/img/staff/inoka_2025.webp)
Chairperson of the Strategic Planning and Management Committee, Faculty of Science- 
Insect pollinators and their role in pollinating watermelon in the Dry Zone of Sri Lanka
Fellow of the Royal
Entomological Society
Fulbright Visiting Scholar Award 2014-2015 – United States - Sri Lanka Fulbright Commission
NSF (National Science Foundation) Merit Awards for Scientific Excellence 2006 in the field of Environment and Biodiversity for the project on ‘Taxonomy and Ecology of Bees of Sri LankMolecular Ecology
Cardoso, G. A., Mariano‐Martins, P., Faria, G. A., Karunaratne, I., Thyssen, P. J., & Torres, T. T. (2025). Divergent Genetic Pathways Underlying Convergent Parasitic Behaviours in Blowflies. Molecular Ecology, 34(11): 1-15.
DOI: https://doi.org/10.1111/mec.17785
Ceylon Journal of Science
Ekanayake, E. M. D., Jayasuriya, K. G., Hemachandra, K. S., Karunarathna, A. K., Athukorala, A. D. S. N. P., Edirisinghe, E. S. M., & Karunaratne, W. A. I. P. (2025). A Case Report on Natural Control of Eggplant Powdery
Mildew Disease by Illeis cincta (Fabricius, 1798) (Coleoptera: Coccinellidae)
in Peradeniya, Sri Lanka . Ceylon Journal of Science, 54(01): 239-245.
DOI: https://doi.org/10.4038/cjs.v54i1.8451
Taprobanica
Gunasekara R.S., Yakandawala K., Jayakody S. Wijesundara D.S.A., Dharmadasa R.M. and Karunaratne W.A.I.P. (2024). Are volatile organic compounds the secret to visitations by pollinators in the food deceptive orchid Vanda tessellata (ROXB.) HOOK. EX G. DON. (Orchidaceae)?. Taprobanica, 13 (02): 39–47.
DOI: https://doi.org/10.47605/tapro.v13i2.329
Journal of Tropical Forest Science
Wijewickrama, T., Karunaratne, I., Wijesundara, S. and Madawala, S. (2022). Impacts of Bambusa bambos spread on seedling recruitment, mortality and regeneration potential of native species in tropical moist evergreen forests of Sri Lanka. . Journal of Tropical Forest Science, 34 (1): 48-62.
Canadian Society of Forensic Science Journal
Bandara W. R. U. A., Karunaratne W. A. I. P., Fuward R. M., Dangolla A. and Yasakeerthi A. D. H. (2022). Estimating
colonization time of maggots infesting wounds in dogs: three case studies. Canadian Society of Forensic Science Journal, .
DOI: 10.1080/00085030.2022.2043521
Ceylon journal of Science
Ekanayake, E.M.D. and Karunaratne, W.A.I.P. (2022). Native Sri Lankan bees enhance pollination of sesame, Sesamum indicum. . Ceylon journal of Science, 51(1): 29-36.
DOI: DOI: http://doi.org/10.4038/cjs.v51i1.7976
Ecological Entomology
Wijewickrama, T., Karunaratne, I., Wijesundara, S. and Madawala, S. (2022). Bambusa bambos (L.) Voss spread influences litter-dwelling
arthropod communities in native forests: A case study from Sri Lanka. . Ecological Entomology, : 1-10.
DOI: 10.1111/een.13182
Journal of Medical Entomology
Bambaradeniya, Y.T.B., Karunaratne, W.A.I.P., Tomberlin J.K. and Magni P.A. (2021). Effect
of type of tissue on the development of Chrysomya rufifacies (Diptera: Calliphoridae) in Sri
Lanka. . Journal of Medical Entomology, 58(4): 1673–1679.
DOI: https://10.1093/jme/tjab044
Journal of Medical Entomology
Lue Cuttiford, Meghan L. Pimsler, Chong Chin Heo, Le Zhend, Inoka Karunaratne, Gabriele Trissini, Aaron M. Tarone, Simonetta Lambiase, Jonathan A. Cammack, and Jeffrey K. Tomberlin (2021). Evaluation of Development datasets for Hermetia illucens (L.) (Diptera:
Stratiomyidae) for estimating the time of placement of human and swine remains in Texas,
USA. Journal of Medical Entomology, 58(4): 1654-1662.
DOI: https://10.1093/jme/tjab081
Ceylon Medical Journal
Gamage P., Anjalie Amarasinghe, Inoka Karunaratne, Rumala Morel, Nalin Nandasiri (2021). Scarabiasis revisited: a case study from Sri Lanka. . Ceylon Medical Journal, 66(1): 53-54.
Wilderness & Environmental Medicine.
Witharana, R.A., Arosha Dissanayake, Inoka Karunaratne, Samantha Wijesinghe (2021). A
rare case of micro-angiopathic hemolytic anemia due to envenoming by giant Asian
honeybee. . Wilderness & Environmental Medicine., S1080-6032(21)00013-2..
DOI: https://10.1016/j.wem.2021.01.008
Journal of Insects as Food and Feed
Perera, K.S.L., Liyanage, C.J., Jinadasa, H.R.N., Gedara, P.K. and Karunaratne, W.A.I.P. (2021). Resource-efficient and eco-friendly model for fruit processing industry waste valorisation using black soldier fly (Hermetia illucens) larvae under tropical conditions. Journal of Insects as Food and Feed, : 1-14.
Ceylon Journal of Science
Galappaththi, M.C.A., Patabendige, N.M., Amarasinghe, S.S., Ranawana, K.B. and Karunaratne, W.A.I.P. (2021). Cochineal Scale Dactylopius opuntiae controls Opuntia dillenii in Bundala National Park, Sri Lanka.
. Ceylon Journal of Science, 50 (03): 297-301.
International Journal of Sustainable Development & World Ecology
Wijewickrama, T., Karunaratne, I., Wijesundara, S. and Madawala, S. (2020). Community perceptions and responses on bamboo spread in native forests: a case study from Sri Lanka. International Journal of Sustainable Development & World Ecology, 27(3): 240-249.
Urban Forestry & Urban Greening
Wijesinghe, E., Minor, E.S., Karunarathne, I. and Yakandawala, K. (2020). Relative attractiveness of ruderals and ornamental plants to flower-visiting insects in a tropical anthropogenic landscape. Urban Forestry & Urban Greening, : 126657.
Ceylon Journal of Science
Wijewickrama, M.P.T., Karunaratne, W.A.I.P., Wijesundara, D.S.A. and Madawala, H.M.S.P. (2020). Bambusa bambos (L.) Voss. Alters. Structure and composition of native forests: A study from moist evergreen forests in Sri Lanka. Ceylon Journal of Science, 49(2): 173-184.
Ceylon Journal of Science
Karunaratne, W.A.I.P. and Karunarathna D.A.G.N.B. (2020). Species assemblages and diversity of litter ants (Hymenoptera: Formicidae) along an altitudinal gradient across two vegetation types in Hanthana Forest in Kandy, Sri Lanka. . Ceylon Journal of Science, 49(3): 317-321.
International Journal of Dermetology
Bambaradeniya, Y.T.B., Karunaratne, W.A.I.P., Rakinawasam, S.V., Tomberlin, J.K., Goonerathne, I. and Kotakadeniya, R.B. (2019). Myiasis incidences reported in and around central
province of Sri Lanka. International Journal of Dermetology, 58: 336–342.
DOI: https://dx.doi.org/10.1111/ijd.14291
Journal of Medical Entomology
Bambaradeniya, Y.T.B., Karunaratne, W.A.I.P., Tomberlin, J.K., Goonerathne, I. and Kotakadeniya, R.B. (2019). Effect of Temperatue and tissue type on the development of myiasis causing fly Chrysomya bezziana (Diptera:
Calliphoridae). Journal of Medical Entomology, 56(3): 625–631.
DOI: https://doi.org/10.1093/jme/tjy212
Journal of Pollination Ecology
Perera, R.A.S.N. and Karunaratne, W.A.I.P. (2019). Floral
visits of the wild bee, Lithurgus atratus
impact yield and seed germinability of okra, Abelmoschus esculentus in Sri Lanka. Journal of Pollination Ecology, 25(1): 1-6.
DOI: https://dx.doi.org/10.26786/1920-7603%282018%2920
International Journal of Fruit Science
Jayarathna, H. L. D. K., Nawarathna, L. S., and Karunaratne, W. A. I. P. (2019). A Novel Approach for Predicting Yield of Seeds in Ficus Fruits Using a Non-Parametric Method. International Journal of Fruit Science, : 1-17.
Ceylon Journal of Science
Perera, P. L. M. M., Jayasuriya, K. M. G. G., Gunaratne, A. M. T. A., Karunaratne, W. A. I. P., Damunupola, J. W., and Prassanna, M. G. M. (2019). Conservation attempt of critically endangered mangrove Lumnitzera littorea (Jack) Voigt in Madu Ganga Ramsar Site of Sri Lanka; stand composition and seed germination. Ceylon Journal of Science, 48(3): 225-234.
Journal of Medical Entomology
Bambaradeniya, Y.T.B., Karunaratne, W.A.I.P., Tomberlin, J.K., Goonerathne, I., Kotakadeniya, R.B. and Magni P.A. (2019). Effect of Temperature and Tissue Type on the Development of the Forensic Fly Chrysomya megacephala (Diptera: Calliphoridae) . Journal of Medical Entomology, 56(6): 1571-1581.
Journal of Medical Entomology
Bambaradeniya, Y.T.B., Karunaratne, W.A.I.P., Tomberlin, J.K., Goonerathne, I. and Kotakadeniya, R.B. (2018). Temperatue and tissue type impact development of Lucilia cuprina (Diptera: Calliphoridae) in Sri Lanka. Journal of Medical Entomology, 55(2): 285–291.
DOI: https://doi.org/10.1093/jme/tjx211
J.Natn.Sci.Foundation Sri Lanka
Silva, T. H. S. E, Diyes, G.C.P, Karunaratne, W.A.I.P. and Edirisinghe, J.P. (2018). Rediscovery of Tetragonula praeterita after 1860: an
unremarked common stingless bee endemic to Sri Lanka. J.Natn.Sci.Foundation Sri Lanka, 46(1): 109-113.
DOI: https://dx.doi.org/10.4038/jnsfsr.v46i1.8271
Sri Lanka Journal of Medicine
Witharana, E.W.R.A., Karunaratne, W.A.I.P., Wijesinghe, S.K.J. and Kadahetti, S.C. L. (2018). Spider bite in
Sri Lanka: Two case reports on envenomation by two Tarantula species. Sri Lanka Journal of Medicine, 27(1): 71-73.
DOI: https://doi.org/10.4038/sljm.v27i1.82
International Journal of Biodiversity
Silva, T. H. S. E., Karunarathna, N. B., and Karunaratne, W. A. I. P. (2017). Competence of Litter Ants for Rapid Biodiversity Assessments. International Journal of Biodiversity, (in press).
Acarologia
Diyes, G.C.P., Karunarathna, D.A.G.N.B., Silva, T.H.S.E., Karunaratne, W. A. I. P. and Rajakaruna, R.S. (2017). Ants as predators of the Spinose Ear Tick, Otobius megnini (Dugès) in Sri Lanka. Acarologia, 57(4): 747-753 (2017).
DOI: https://doi.org/10.1051/acarologia/20174200
Psyche
Jayasinghe S., Silva S. E. and Karunaratne W.A.I.P. (2017). Buzzing wild bee visits enhance seed set in eggplant, Solanum melongena. Hindawi Publishing Corporation. Psyche, Volume 2017: Article ID 4624062, 7 pages.
DOI: https://doi.org/10.1155/2017/4624062
Journal of the National Science Foundation
Karunaratnea, W.A.I.P. , Edirisinghe, J.P. and Engel, Michael S. (2017). First record of a tear-drinking stingless bee, Lisotrigona cacciae (Nurse) (Hymenoptera: Apidae: Meliponini), from the central hills of Sri Lanka. Journal of the National Science Foundation, 45 (1): 79-81.
DOI: https://dx.doi.org/10.4038/jnsfsr.v45i1.8042
International Journal of Tropical Insect Science
Abeyasuriya, K.G.T.N., Nugapola, N.W.N.P., Perera, M.D.B., Karunaratne, W. A. I. P. , and Karunaratne S.H.P.P. (2017). Effect of dengue mosquitoe control insecticide fogging on non-target insects. International Journal of Tropical Insect Science, 37(1): 11-18 .